I only have SCAG.
While it doesn't specify on the type of tribute to the goddess, I believe that it should be something that would feed the little critters. :) (correct me if I'm wrong, but I personally believe that to be a really cute tribute.)
There's also a note about the area behind her shrine being a no hunt/scavage. So I can guess that it's a pretty safe interpretation that she is indeed against unnecessary savaging, hence Davros looks down on it too.
OOC:
I'm sorry if it bothers you, but I usually like discussing things like these with the GM, so that we're on the same page.